THE ELEMENTS OF COMPLETE PLANT NUTRITION
The following is a brief guideline of the role of essential and beneficial mineral nutrients that are crucial for growth. Eliminate any one of these elements, and plants will display abnormalities of growth, deficiency symptoms, or may not reproduce normally.
Iron is necessary for many enzyme functions and as a catalyst for the synthesis of chlorophyll. It is essential for the young growing parts of plants. Deficiencies are pale leaf colour of young leaves followed by yellowing of leaves and large veins. Iron is lost by leaching and is held in the lower portions of the soil structure. Under conditions of high pH (alkaline) iron is rendered unavailable to plants. When soils are alkaline, iron may be abundant but unavailable. Applications of an acid nutrient formula containing iron chelates, held in soluble form, should correct the problem.
Manganese is involved in enzyme activity for photosynthesis, respiration, and nitrogen metabolism. Deficiency in young leaves may show a network of green veins on a light green background similar to an iron deficiency. In the advanced stages the light green parts become white, and leaves are shed. Brownish, black, or greyish spots may appear next to the veins. In neutral or alkaline soils plants often show deficiency symptoms. In highly acid soils, manganese may be available to the extent that it results in toxicity.
Boron is necessary for cell wall formation, membrane integrity, and calcium uptake and may aid in the translocation of sugars. Boron affects at least 16 functions in plants. These functions include flowering, pollen germination, fruiting, cell division, water relationships and the movement of hormones. Boron must be available throughout the life of the plant. It is not translocated and is easily leached from soils. Deficiencies kill terminal buds leaving a rosette effect on the plant. Leaves are thick, curled and brittle. Fruits, tubers and roots are discoloured, cracked and flecked with brown spots.
Zinc is a component of enzymes or a functional cofactor of a large number of enzymes including auxins (plant growth hormones). It is essential to carbohydrate metabolism; protein synthesis and inter nodular elongation (stem growth). Deficient plants have mottled leaves with irregular chlorotic areas. Zinc deficiency leads to iron deficiency causing similar symptoms. Deficiency occurs on eroded soils and is least available at a pH range of 5.5 - 7.0. Lowering the pH can render zinc more available to the point of toxicity.
Copper is concentrated in roots of plants and plays a part in nitrogen metabolism. It is a component of several enzymes and may be part of the enzyme systems that use carbohydrates and proteins. Deficiencies cause die back of the shoot tips, and terminal leaves develop brown spots. Copper is bound tightly in organic matter and may be deficient in highly organic soils. It is very mobile and used with crop growing and lost from the soil.
Molybdenum is a structural component of the enzyme that reduces nitrates to ammonia. Without it, the synthesis of proteins is blocked and plant growth ceases. Root nodule (nitrogen fixing) bacteria also require it. Seeds may not form completely, and nitrogen deficiency may occur if plants are lacking molybdenum. Deficiency signs are pale green leaves with rolled or cupped margins.
Chlorine is involved in osmosis (movement of water or solutes in cells), the ionic balance necessary for plants to take up mineral elements and in photosynthesis. Deficiency symptoms include wilting, stubby roots, chlorotic (yellowing) and bronzing. Odours in some plants may be decreased. Chloride, the ionic form of chlorine used by plants, is usually found in soluble forms and is lost by leaching. Some plants may show signs of toxicity if levels are too high.
Nickel has just recently won the status as an essential trace element for plants according to the Agricultural Research Service Plant, Soil and Nutrition Laboratory in Ithaca, NY. It is required for the enzyme urease to break down urea to liberate the nitrogen into a usable form for plants. Nickel is required for iron absorption. Seeds need nickel in order to germinate. Plants grown without additional nickel will gradually reach a deficient level at about the time they mature and begin reproductive growth. If nickel is deficient plants may fail to produce viable seeds.
Sodium is involved in osmotic (water movement) and ionic balance in plants.
Cobalt is required for nitrogen fixation in legumes and in root nodules of non legumes. The demand for cobalt is much higher for nitrogen fixation than for ammonium nutrition. Deficient levels could result in nitrogen deficiency symptoms.
Silicon is found as a component of cell walls. Plants with supplies of soluble silicon produce stronger, tougher cell walls making them a mechanical barrier to piercing and sucking insects. This significantly enhances plant heat and drought tolerance. Foliar sprays of silicon have also shown benefits reducing populations of aphids on field crops. Tests have also found that silicon can be deposited by the plants at the site of infection by fungus to combat the penetration of the cell walls by the attacking fungus. Improved leaf erectness, stem strength and prevention or depression of iron and manganese toxicity has all been noted as effects from silicon. Silicon has not been determined essential for all plants but may be beneficial for many.
Nitrogen is a major component of proteins, hormones, chlorophyll, vitamins and enzymes essential for plant life. Nitrogen metabolism is a major factor in stem and leaf growth (vegetative growth). Too much can delay flowering and fruiting. Deficiencies can reduce yields, because yellowing of the leaves and stunt growth.
Phosphorus is necessary for seed germination, photosynthesis, protein formation and almost all aspects of growth and metabolism in plants. It is essential for flower and fruit formation. Low pH (<4) results in phosphate being chemically locked up in organic soils. Deficiency symptoms are purple stems and leaves; maturity and growth are retarded. Yields of fruit and flowers are poor. Premature drop of fruits and flowers may often occur. Phosphorus must be applied close to the plant's roots in order for the plant to utilize it. Large applications of phosphorus without adequate levels of zinc can cause a zinc deficiency.
Potassium is necessary for formation of sugars, starches, carbohydrates, protein synthesis and cell division in roots and other parts of the plant. It helps to adjust water balance, improves stem rigidity and cold hardiness, enhances flavour and colour on fruit and vegetable crops, increases the oil content of fruits and is important for leafy crops. Deficiencies result in low yields, mottled, spotted or curled leaves, scorched or burned look to leaves.
Sulphur is a structural component of amino acids, proteins, vitamins and enzymes and is essential to produce chlorophyll. It imparts flavour to many vegetables. Deficiencies show as light green leaves. Sulphur is readily lost by leaching from soils and should be applied with a nutrient formula.
Magnesium is a critical structural component of the chlorophyll molecule and is necessary for functioning of plant enzymes to produce carbohydrates, sugars and fats. It is used for fruit and nut formation and essential for germination of seeds. Deficient plants appear chlorotic, show yellowing between veins of older leaves; leaves may drop. Magnesium is leached by watering and must be supplied when feeding.
Calcium activates enzymes, is a structural component of cell walls, influences water movement in cells and is necessary for cell growth and division. Some plants must have calcium to take up nitrogen and other minerals. Calcium is easily leached. Calcium, once deposited in plant tissue, is immobile (non-translocatable) so there must be a constant supply for growth. Deficiency causes stunting of new growth in stems, flowers and roots. Symptoms range from distorted new growth to black spots on leaves and fruit. Yellow leaf margins may also appear.
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Rare Earths (REE)
A recent update of the mineral composition of the Elverdton copper mine resource revealed the Rare Earth Minerals (REE) Lanthanum La and Cerium Ce to be present at levels to advantage crops at risk from moisture stress. As outlined in research carried out by Rural Industries and Development Corporation, Impact of Rare Earth Elements on plant physiology productivity.
Publication Number 01/45
Project Number DAV – 122A